199 species of dung beetles are known to occur in Angloa XXX of which are endemic. If you are aware of other species not included here please e-mail c.borrow@nhm.ac.uk or write to Chris Borrow, Department of Entomology, The Natural History Museum, Cromwell Road, London, England, SW7 5BD.
- ANACHALCOS
- Anachalcos convexus Boheman 1857 (BMNH drawer 63)
- Anachalcos cupreus Fabricius 1775 (BMNH drawer 63)
- Anachalcos procerus Gerstaecker 1874 (BMNH drawer 64)
- CACCOPHILUS
- Caccophilus semiaeneus d'Orbigny 1905 (BMNH drawer 202)
- CATHARSIUS
- Catharsius alpheus Gillet 1933 (BMNH drawer 116)
- Catharsius bellus Gillet 1918
- Catharsius biconifer Felsche 1907 (BMNH drawer 117)
- Catharsius cassius Kolbe 1893
- Catharsius dux Harold 1878 (BMNH drawer 113)
- Catharsius fastidiosus Thomson 1858 (BMNH drawer 118)
- Catharsius gorilloides Felsche 1907
- Catharsius machadoi Ferreira 1960
- Catharsius melancholicus Boheman 1860 (BMNH drawer 116)
- Catharsius pandion Harold 1877
- Catharsius satyrus Kolbe 1893 (BMNH drawer 114)
- Catharsius sesostris Waterhouse 1888 (BMNH drawer 118)
- Catharsius tricornutus Degeer 1778 (BMNH drawer 117)
- COPRIDASPIDUS
- Copridaspidus peregrinus Harold 1878 (BMNH drawer 142)
- COPRIS
- Copris bovinus Gillet 1908 (BMNH drawer 137)
- Copris draco Arrow 1906 (BMNH drawer 132)
- Copris elphenor Klug 1855 (BMNH drawer 131)
- Copris inhalatus Quedenfeldt 1884 (BMNH drawer 132)
- Copris macer Peringuey 1901 (BMNH drawer 137)
- Copris mesacanthus Harold 1878 (BMNH drawer 133)
- Copris orion Klug 1835 (BMNH drawer 136)
- Copris orphanus Guerin 1847 (BMNH drawer 137)
- Copris vilhenai Ferreira 1962
- COPTORHINA
- Coptorhina angolensis Arrow 1906 (BMNH drawer 71)
- Coptorhina auspicata Peringuey 1901 (BMNH drawer 71)
- DELOPLEURUS
- Delopleurus pullus Boheman 1857 (BMNH drawer 69)
- DIASTELLOPALPUS
- Diastellopalpus infernalis Lansberge 1887 (BMNH drawer 209)
- Diastellopalpus lamellicollis Quedenfeldt 1884 (BMNH drawer 209)
- Diastellopalpus tridens Fabricius 1781 (BMNH drawer 209)
- DIGITONTHOPHAGUS
- Digitonthophagus gazella Fabricius 1787 (BMNH drawer 221)
- DREPANOCERUS
- Drepanocerus kirbyi Kirby 1828 (BMNH drawer 186)
- Drepanocerus laticollis Fahraeus 1857 (BMNH drawer 186)
- EUONITICELLUS
- Euoniticellus nasicornis Reiche 1849 (BMNH drawer 190)
- Euoniticellus triangulatus Harold 1873 (BMNH drawer 190)
- Euonthophagus carbonarius Klug 1855 (BMNH drawer 220)
- FURCONTHOPHAGUS
- Furconthophagus variegatus Fabricius 1798 (BMNH drawer 231)
- GARRETA
- Garreta caffer Fahraeus 1857 (BMNH drawer 31)
- Garreta nitens Olivier 1789 (BMNH drawer 33.1)
- GYMNOPLEURUS
- Gymnopleurus aenescens Wiedemann 1821 (BMNH drawer 36 & 38.3)
- Gymnopleurus ignitus Klug 1855 (BMNH drawer 38)
- Gymnopleurus sericatus Erichson 1843 (BMNH drawer 38)
- Gymnopleurus virens Erichson 1843 (BMNH drawer 37)
- HELIOCOPRIS
- Heliocopris andersoni Bates 1868
- Heliocopris antenor Olivier 1789 (BMNH drawer 86)
- Heliocopris atropos Boheman 1860 (BMNH drawer 86)
- Heliocopris fonsecai Ferreira 1967
- Heliocopris haroldi Kolbe 1893 (BMNH drawer 84)
- Heliocopris hunteri Waterhouse 1891 (BMNH drawer 87)
- Heliocopris japetus Klug 1855 (BMNH drawer 80)
- Heliocopris kolbei Felsche 1901 (BMNH drawer 84)
- Heliocopris pauliani Janssens 1940
- Heliocopris primal Fabricius 1798 (BMNH drawer 83)
- Heliocopris samson Harold 1878
- KHEPER
- Kheper cuvieri Macleay 1821 (BMNH drawer 22)
- Kheper festivus Harold 1868 (BMNH drawer 20)
- Kheper intermedius Gillet 1909 (BMNH drawer 20)
- Kheper lamarcki Macleay 1821 (BMNH drawer 21)
- Kheper prodigiosus Erichson 1843 (BMNH drawer 21)
- Kheper subaeneus Harold 1869 (BMNH drawer 22)
- LIATONGUS
- Liatongus interruptus Quedenfeldt 1884 (BMNH drawer 193)
- Liatongus militaris Castelnau 1840 (BMNH drawer 194)
- Liatongus testudo Lansberge 1886 (BMNH drawer 194)
- METACATHARSIUS
- Metacatharsius dentinum Ferreira 1964 (BMNH drawer 121)
- Metacatharsius exiguus Boheman 1860 (BMNH drawer 121)
- Metacatharsius ferreirae Ferreira 1962
- Metacatharsius opacus Waterhouse 1891 (BMNH drawer 120)
- Metacatharsius troglodytes Boheman 1857 (BMNH drawer 121)
- MILICHUS
- Milichus apicalis Fahraeus 1857 (BMNH drawer 204)
- NEOSISYPHUS
- Neosisyphus rugosus Gory 1833 (BMNH drawer 42.3)
- ONITICELLUS
- Oniticellus formosus Chevrolat 1830 (BMNH drawer 191)
- Oniticellus planatus Castelnau 1840 (BMNH drawer 191)
- ONITIS
- Onitis abyssinicus Reiche 1847
- Onitis aerarius Harold 1878 (BMNH drawer 184)
- Onitis alexis Klug 1835 (BMNH drawer 181)
- Onitis cupreus Castelnau 1840 (BMNH drawer 184)
- Onitis curvipes Lansberge 1875 (BMNH drawer 179)
- Onitis deceptor Peringuey 1901 (BMNH drawer 179)
- Onitis fabricii Roth 1851 (BMNH drawer179)
- Onitis obscurus Lansberge 1886 (BMNH drawer184)
- Onitis orthopus Lansberge 1875
- Onitis setosus Lansberge 1875 (BMNH drawer184)
- Onitis sphinx Fabricius 1775 (BMNH drawer181)
- Onitis vanderkelleni Lansberge 1886 (BMNH drawer183)
- ONTHOPHAGUS
- Onthophagus africanus Lansberge 1886
- Onthophagus albipodex d'Orbigny 1902 (BMNH drawer268)
- Onthophagus angolensis Paulian 1937
- Onthophagus anomalipes d'Orbigny 1902 (BMNH drawer256)
- Onthophagus apiciosus d'Orbigny 1902 (BMNH drawer266)
- Onthophagus axillaris Boheman 1860
- Onthophagus benguelianus Paulian 1937
- Onthophagus bicarinatus Gomes & Alves 1944
- Onthophagus bidens Olivier 1789 (BMNH drawer269)
- Onthophagus bubalus Harold 1867 (BMNH drawer264)
- Onthophagus cancer Lansberge 1886 (BMNH drawer268)
- Onthophagus cinctipennis Quedenfeldt 1884 (BMNH drawer256)
- Onthophagus clitellarius d'Orbigny 1908
- Onthophagus colasi Paulian 1937
- Onthophagus cornifrons Thomson 1858 (BMNH drawer258)
- Onthophagus dispar Peringuey 1901 (BMNH drawer264)
- Onthophagus duvivieri d'Orbigny 1904 (BMNH drawer263)
- Onthophagus fallax d'Orbigny 1913 (BMNH drawer273)
- Onthophagus fimetarius Roth 1851 (BMNH drawer268)
- Onthophagus foliaceus Lansberge 1886 (BMNH drawer265)
- Onthophagus foliiceps Quedenfeldt 1884
- Onthophagus furcatoides Lansberge 1886
- Onthophagus furculifer d'Orbigny 1905 (BMNH drawer273)
- For Onthophagus gazella see Digitonthophagus gazella Fabricius, 1787 (BMNH drawer 221)
- Onthophagus gibbus d'Orbigny 1913 (BMNH drawer265)
- Onthophagus gonopygus d'Orbigny 1902 (BMNH drawer270)
- Onthophagus grandidorsis d'Orbigny 1913
- Onthophagus granulifer Harold 1886 (BMNH drawer270)
- Onthophagus granulipennis Lansberge 1886 (BMNH drawer258)
- Onthophagus guttatus Boheman 1860
- Onthophagus humpatensis Lansberge 1886 (BMNH drawer268)
- Onthophagus illotus Peringuey 1900 (BMNH drawer257)
- Onthophagus importunus Peringuey 1901 (BMNH drawer256)
- Onthophagus impressicollis Boheman 1860 (BMNH drawer268)
- Onthophagus insignis Peringuey 1896 (BMNH drawer268)
- Onthophagus jugicola d'Orbigny 1902 (BMNH drawer262)
- Onthophagus laeviceps d'Orbigny 1902 (BMNH drawer267)
- Onthophagus lamelliger Gerstaecker 1871 (BMNH drawer270)
- Onthophagus laminicornis Lansberge 1886
- Onthophagus monardiellus Frey 1963
- Onthophagus monradi Boucomont 1936
- Onthophagus nanus Harold 1878 (BMNH drawer259)
- Onthophagus ochropygus d'Orbigny 1902 (BMNH drawer272)
- Onthophagus parcenotatus d'Orbigny 1913
- Onthophagus parumnotatus Fahraeus 1857 (BMNH drawer263)
- Onthophagus paucigranosus Lansberge 1886 (BMNH drawer265)
- Onthophagus placens Peringuey 1904
- Onthophagus politissimus d'Orbigny 1908 (BMNH drawer259)
- Onthophagus probus Peringuey 1900 (BMNH drawer267)
- Onthophagus proximus d'Orbigny 1908 (BMNH drawer268)
- Onthophagus pugionatus Fahraeus 1857
- Onthophagus quadrinodosus Fahraeus 1857 (BMNH drawer264)
- Onthophagus rhinocerus Gomes-Alves 1944 (BMNH drawer274)
- Onthophagus rubricatus d'Orbigny 1902 (BMNH drawer266)
- Onthophagus rufonotatus d'Orbigny 1902
- Onthophagus sansibaricus Harold 1878 (BMNH drawer263)
- Onthophagus schaufussi Harold 1867 (BMNH drawer269)
- Onthophagus sellatus Klug 1845 (BMNH drawer231)
- Onthophagus semiflavus Boheman 1860 (BMNH drawer259)
- Onthophagus seminitens d'Orbigny 1902
- Onthophagus semivestitus Paulian 1937
- Onthophagus setosus Fahraeus 1857 (BMNH drawer262)
- Onthophagus simplex Raffray 1877 (BMNH drawer262)
- Onthophagus stellio Erichson 1843 (BMNH drawer272)
- Onthophagus suffusus Klug 1855 (BMNH drawer272)
- Onthophagus sumptuosus Paulian 1937
- Onthophagus testaceoviolaceus Paulian 1937
- Onthophagus tripartitus d'Orbigny 1902 (BMNH drawer267)
- Onthophagus variegranosus d'Orbigny 1905 (BMNH drawer263)
- Onthophagus vesanus Balthasar 1967
- Onthophagus vinctus Erichson 1843 (BMNH drawer258)
- PACHYLOMERUS
- Pachylomerus femoralis Kirby 1828 (BMNH drawer24)
- PARAPHYTUS
- Paraphytus africanus Boucomont 1923 (BMNH drawer69)
- PEDARIA
- Pedaria angolana Boucomont 1922
- Pedaria criberrima Waterhouse 1890 (BMNH drawer72)
- Pedaria cylindrica Fahraeus 1857 (BMNH drawer72)
- Pedaria granulosa Ferreira 1962
- Pedaria ovata Boucomont 1922
- Pedaria rohani Boucomont 1922
- Pedaria segregis Peringuey 1901 (BMNH drawer72)
- Pedaria spinosa Boucomont 1922 (BMNH drawer72)
- Pedaria tuberculigera Waterhouse 1890 (BMNH drawer72)
- PHALOPS
- Phalops boschas Klug 1855 (BMNH drawer208)
- Phalops prasinus Erichson 1843 (BMNH drawer208)
- Phalops rufatus d'Orbigny 1902
- PROGODERUS
- Proagoderus bicallosus Klug 1855 (BMNH drawer217)
- Proagoderus brucei Reiche 1849 (BMNH drawer218)
- Proagoderus panoplus Bates 1888 (BMNH drawer215)
- Proagoderus rasidorsis d'Orbigny 1905 (BMNH drawer218)
- Proagoderus sapphirinus Fahraeus 1857 (BMNH drawer217)
- Proagoderus speculicollis Quedenfeldt 1884 (BMNH drawer217)
- Proagoderus subextensus Kolbe 1895 (BMNH drawer216)
- Proagoderus superbus d'Orbigny 1904
- Proagoderus trianguliceps d'Orbigny 1905 (BMNH drawer218)
- Proagoderus unidens Quedenfeldt 1884 (BMNH drawer249)
- PSEUDOCOPRIS
- Pseudocopris lunaticeps Gillet 1910
- PSEUDOSAPROECIUS
- Pseudosaproecius cylindroides d'Orbigny 1908 (BMNH drawer70)
- Pseudosaproecius validicornis Quedenfeldt 1884 (BMNH drawer70)
- SCARABAEOLUS
- Scarabaeolus flavicornis Boheman 1860 (BMNH drawer1)
- Scarabaeolus knobeli Ferreira 1958 (BMNH drawer2)
- SCARABAEOLUS
- Scarabaeus cancer Arrow 1919 (BMNH drawer27)
- Scarabaeus gilleti Janssens 1940 (BMNH drawer2)
- Scarabaeus obsoletepunctatus Balthasar 1940 (BMNH drawer1)
- Scarabaeus palemo Olivier 1789 (BMNH drawer1)
- Scarabaeus plausibilis Peringuey 1892 (BMNH drawer8)
- Scarabaeus poggei Waterhouse 1890 (BMNH drawer8)
- Scarabaeus sulcipennis Quedenfeldt 1888 (BMNH drawer8)
- Scarabaeus zambezianus Peringuey 1901 (BMNH drawer9)
- SCELIAGES
- Sceliages augias Gillet 1908 (BMNH drawer27.1)
- SISYPHUS
- Sisyphus biarmatus Felsche 1909 (BMNH drawer41)
- TINIOCELLUS
- Tiniocellus spinipes Roth 1851 (BMNH drawer190)
- TOMOGONUS
- Tomogonus crassus d'Orbigny 1904




